Reproduction · Process
Fertilisation, seeds and fruit
Pollen has landed. The two gametes are still several centimetres apart, and neither of them can move.
Start here
A pollen grain solves the problem by growing a tunnel.
There is no sperm here, nothing swims, and nothing is carried. The gametes are at opposite ends of a solid stalk of plant tissue, and the plant has to get one of them to the other.
So how does the male gamete reach the ovule?
The grain grows. Once it lands on a stigma of the right species it puts out a pollen tube — a single cell extending down through the tissue of the style, digesting its way towards an ovary it cannot see — and the male gamete nucleus travels down inside it. The journey takes hours in some plants and months in others. Only when the tube reaches an ovule does anything fuse, and that is the moment this lesson is about.
Before and after · six parts
What each part turns into
0 of 6 opened
A fertilised flower does not build a fruit from nothing — it converts the parts it already has. Tap a row to see why.
Why: One ovule becomes one seed, which is why counting the seeds in a fruit counts the ovules that were fertilised. The seed contains an embryo plant, a food store, and a tough coat around both.
Why: It swells, and what it swells into depends on the species: fleshy in a tomato or a plum, dry and papery in a pea pod, hard in a hazelnut. All of them are ovaries doing the same job — protecting the seeds and getting them moved.
Why: This is the fertilised cell, and it divides to become a miniature plant with the beginnings of a root and a shoot, folded up inside the seed and waiting. It is the only part of all this that is genuinely new.
Why: The advertising budget is cut the moment the sale is made. A flower whose ovules are fertilised has nothing to gain from further insect visits, so the petals — which were expensive to build — are abandoned.
Why: Same reasoning. The male parts of this flower have already sent their pollen out into the world, and it is other flowers they have to reach, not this one.
Why: Look at the top of a strawberry or a tomato: that little green star is the sepals, still there long after everything else has gone. They protected the bud and in many species they go on protecting the fruit.
The process, in order
Five steps, and only one of them is fertilisation.
Pollen lands · On the stigma — this is pollination
A grain arrives, carried by an insect or the wind. If it is the wrong species the stigma usually does not respond at all; if it is the right one, the grain begins to grow.
The pollen tube grows · Down through the style
The grain extends a single tube down through the tissue of the style towards the ovary, digesting its way through as it goes. The male gamete nucleus travels down inside it.
Fertilisation · Inside an ovule — and nowhere else
The tube reaches an ovule and the male gamete nucleus fuses with the female gamete nucleus. That fusion, and only that fusion, is fertilisation. Every ovule that is going to become a seed needs its own pollen tube.
Seeds form · From the fertilised ovules
Each fertilised ovule becomes a seed: an embryo plant, a store of food to start it off, and a tough coat around both. The seed can survive conditions that would kill the parent plant.
The fruit forms · From the ovary around them
The ovary wall swells and becomes the fruit, and the rest of the flower withers away. Whether it ends up fleshy, dry, hooked or winged depends on how that species intends to move its seeds — which is the next lesson.
Key fact
Pollination is delivery to the stigma. Fertilisation is fusion at the ovule, later, at the bottom of a tube the pollen grain had to grow. A fertilised ovule becomes a seed and the ovary around it becomes the fruit.
Think again
“Pollination and fertilisation are the same thing.”
They are separated by a distance, a delay and a definition — and this is the single most common error in the whole of plant reproduction. Pollination is transport: a grain of pollen arrives on a stigma. Nothing has joined, nothing is fertilised, and the plant may still end up with no seeds at all. Fertilisation is the fusion of the pollen's gamete nucleus with the ovule's, which happens somewhere else entirely — down inside the ovary — after the grain has grown a tube the whole length of the style to get there. Hours later in some species; in an oak, months. It is the same distinction you drew in Gametes and fertilisation between a sperm reaching an egg and a sperm fusing with one. Arriving is not fusing, in an animal or in a plant.
“A tomato is a vegetable.”
In a kitchen, certainly. In biology the word fruit has a definition and it has nothing to do with sweetness: a fruit is the structure that develops from the ovary of a flower, and it contains the seeds. Cut anything open and the seeds settle it. Tomatoes, cucumbers, peppers, courgettes, pea pods, aubergines and — the one that surprises everybody — the tough case around a grain of wheat are all fruits. Carrots are roots, potatoes are underground stems, celery is a leaf stalk, and none of those has ever been near an ovary. Vegetable is a culinary word covering any plant part we happen to eat savoury; it draws no biological line at all. This is worth having straight for a reason beyond winning an argument: once you know a fruit is a converted ovary, you can look at any one and read backwards to the flower it used to be.
Mastery ladder
Not started yet.
Rungs 3 and 4 you mark yourself.
Rung 1 · Define it precisely
In a flowering plant, which of these is fertilisation?
Rung 2 · The one that catches people
Which of these is a fruit, biologically?
Rung 3 · Explain the sequence
Describe what happens from a pollen grain landing on a stigma to a fruit forming. Make clear which single step is fertilisation, and say what the ovule and the ovary each become.
Rung 4 · Take it somewhere new
A gardener pinches off every flower on a tomato plant as soon as it opens, to keep the plant tidy. Explain why they will get no tomatoes, using the words pollination, fertilisation, ovule and ovary.
Key note
After pollination, the pollen grain grows a pollen tube down through the style to an ovule. The male gamete nucleus travels down the tube and fuses with the female gamete nucleus inside the ovule: that fusion is fertilisation. The fertilised ovule becomes a seed containing an embryo plant and a food store, inside a tough coat. The ovary around it becomes the fruit. Petals, stamens and stigma have done their jobs and wither.
Going further
Flowering plants fertilise twice, with the same pollen tube. Two nuclei travel down it: one fuses with the egg nucleus and becomes the embryo, and the second fuses with other nuclei in the ovule to make a tissue called endosperm — the seed's food store. Nothing else on Earth does this, and it solves an accounting problem elegantly. A food store is expensive, so a plant that built one for every ovule would waste most of them; here the store is only started once fertilisation has actually happened, and an unfertilised ovule costs almost nothing. Then consider what that tissue is. A grain of wheat, rice, maize, barley or oats is mostly endosperm; white flour is very nearly pure endosperm with the embryo and the coat milled away. Most of the calories eaten by most people who have ever lived have been a plant's packed lunch for a seedling, redirected — and none of it would have been made at all if a pollen tube had not arrived.
Before this lesson
- This follows on from Flowers and pollination.
Connects to
At GCSE this becomes
- Seed structure and germination, plant hormones, and the genetics of self- and cross-fertilisation.
Where to next
Ask Mr Badmus AI
Want to check the difference between pollination and fertilisation?
Timings vary enormously between species: a pollen tube may reach the ovule in a few hours or, in some trees, after several months.
Lesson content © MrBadmusAI.